Introduction
a thymidine analog that incorporates DNA of
dividing cells during the S-phase of the cell cycle. However,
BrdU is not a marker of the S-phase of the cell cycle. As a
thymidine analog, it is a marker of DNA synthesis.
PMID: 17020783
an exogenously administered DNA precursor
label and used as a proliferation marker
PMID: 7912188,
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BrdU Incorporation Techniques
BrdU can be administered to laboratory
animals via IP injections, is readily incorporated into nuclei
during the DNA synthetic phase of the cell cycle, and is
detected with an anti-BrdU antibody.
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BrdU was instilled intraperitoneally and the
animals were painlessly sacrificed between 1 hour and 10 days
after the injection.
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both dose and exposure time to BrdU may
influence the final results when cell proliferation is assessed,
the variations obtained clearly depending on the technique used
for the immunological detection of BrdU-positive S-phase cells.
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Each patient received a local injection of 300 mg to 500 mg of BrdU into the cervix, at the time of endocervical laser conization, to label S-phase cells in neoplastic tissue. Labeled cells in normal and neoplastic tissues were detected in biopsy specimens or smears by indirect immunoperoxidase staining using the anti-BrdU MAb. PMID: 2350389
Specimens were incubated in vitro with
BrdUrd and then fixed and paraffin embedded. Sections were
immunohistochemically stained with antibodies to BrdUrd,
proliferating cell nuclear antigen (PCNA), and Ki-67.
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Comparison
Comparison of cell proliferation index in
equine and caprine embryos using a modified BrdU incorporation
assay.
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both ISH for histone mRNA and IHC with Ki-67
(MIB-5) are preferable techniques for assessment of cell
proliferation in rat paraffin-embedded renewing tissues compared
to PCNA IHC. They can substitute for BrdU IHC when necessary.
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The BrdU and cyclin A indices were
significantly correlated with each other. In the more dysplastic
cases, the cyclin A LI was quantitatively much larger than that
for BrdU, suggesting that the protein was being overexpressed.
It was concluded that as a tool to study the kinetic aspects of
the cell cycle in dysplastic lesions cyclin A was limited by the
fact that it is overexpressed. In minimally dysplastic lesions
and normal epithelia, however, cyclin A may be a viable
alternative to BrdU for the study of the S-phase.
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PCNA has a lower variation between biopsies
than BrdU, but higher variation between scorings. When used in a
clinical or epidemiological setting, it is important to take
multiple biopsies at multiple time points.
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MIB-1 is closely associated with BrdU in
clinicopathologic findings and is a more useful tool for
evaluating cell proliferation than PCNA.
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The labelling indices determined by histone
NISH and BrdU incorporation were similar, whereas that of Ki67
IHC was four times greater.
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proliferation rate in livers of 120 mice (60
males and 60 females) was analyzed by immunohistochemical
detection of bromodeoxyuridine (BrdU) incorporation and
proliferating cell nuclear antigen (PCNA) expression on
ethanol-fixed/paraffin-embedded specimens.
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Double label immunofluorescence against PCNA
and BrdU clearly revealed several characteristics of DNA
replication sites in synchronized CHO cells.
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PCNA could replace the BrdU method for
identifying the proliferating cells, and the major advantages of
PCNA method is that it could be done without any pretreatment
and avoid injection of the teratogenic agent for diagnostic
purpose.
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percentage of PCNA-positive cells was correlated with the BrdU labelling index and the histological malignancy of the brain tumours. The correlation coefficient was 0.84. This suggests that the immunohistochemical staining for PCNA in paraffin sections is a good alternative to the BrdU labelling index. PMID: 1357920
Expression
BrdU is a reliable standard and a more useful
tool for the evaluation of proliferative activity of breast
tumors.
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a significant increase in BrdU-positive cells
in the neonatal mouse hippocampus in the injured area compared
to the non-injured area, most prominent in the dentate gyrus
(DG).
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Increases in BrdU-positive cells were
detected in the hippocampal dentate gyrus at 5, 7, and 9 days
after ischemia.
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Function
BrdU application does not induce apoptosis or
necrosis as revealed with the annexin V/PI assay. We concluded
that continuous BrdU treatment did not affect cell viability,
but essentially alters cell cycle progression in three out of
four cell lines tested.
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BrdU labeling in combination with
histopathological observation is therefore a reliable approach
to assessment of test compound effects in vivo.
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Incorporation of BrdU into the nuclei during
a period of 4 h was used to determine the proliferation of the
embryonic cells in vitro.
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Applications
Electron Microscopy (EM)
A simplified method of immunostaining of
bromodeoxyuridine (BrdU), a thymidine analogue, for routinely
paraformaldehyde or glutaraldehyde and osmium tetroxide-fixed
and epon embedded specimens was established using regenerating
rat liver.
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Demonstration of chromosome replication by
BrdU antibody technique and electron microscopy.
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ELISA
An ELISA-based method for the quantification
of incorporated BrdU as a measure of cell proliferation in vivo.
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a simple, sensitive, nonradioactive,
relatively rapid and relatively inexpensive protocol to measure
DNA synthesis in cultured cells by a chemiluminescent
bromodeoxyuridine (BrdU) enzyme-linked immunosorbent assay
(ELISA).
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BrdU ELISA is the most sensitive of the three
proliferation assays used for the assessment of CD4 + T
lymphocyte growth and is the preferred assay when small changes
in cell growth are expected.
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Flow Cytometry (FC)
Three cell preparation protocols for BrdU
measurement were compared for their ability to maintain
fluorescent surface staining and scatter parameters of in vivo
BrdU-labeled cells by flow cytometry.
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bromodeoxyuridine (BrdU) incorporation assay
by flow cytometry was compared with 3H-TdR incorporation assay.
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cell kinetics were examined by flow cytometry with BrdU-PI double staining. PMID: 1284799
Flow cytometric bromodeoxyuridine (BrdU)/DNA
analysis using fresh solid tumors and its clinical significance.
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Bivariate distributions of BrdU and DNA
content are simultaneously obtained by flow cytometric two
parameter analysis with a double staining method.
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Studies on antitumor effects and mechanism of
action of l-hexylcarbamoyl-5-fluorouracil by DNA/BrdU double
staining using flow cytometry.
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a method for the estimation of the
proliferative activity of tumour cells obtained by fine needle
sampling without aspiration (FNS), using simultaneously S-phase
fractions (SPF) measured on DNA histograms and
5-bromodeoxyuridine (BrdU) labelling index (BLI) measured by
flow cytometry.
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BrdU-Hoechst flow cytometry: a unique tool
for quantitative cell cycle analysis.
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Peripheral blood lymphocyte cultures of a
patient with clinical evidence of Fanconi's anemia (FA) were
investigated by means of BrdU/Hoechst flow cytometry.
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Differences in growth factor sensitivity
between primary and transformed murine cell cultures revealed by
BrdU/Hoechst flow cytometry.
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An improved BrdU-Hoechst flow assay was
applied to cell kinetic studies of human lymphocyte cultures
during a 24-96 hr interval after PHA stimulation.
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Cell cycle kinetics by BrdU-Hoechst flow
cytometry: an alternative to the differential metaphase
labelling technique.
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Flow cytometric analysis of chromosomes and
cells using a modified BrdU-Hoechst method.
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Immunocytochemistry
Cell proliferation was assessed immunocytochemically using BrdU incorporation technique and documented on both ganglionic sections and microglia cultured cells at different experimental conditions and times after activation. PMID: 15270245
Cells in S-phase of cell cycle were labelled
in vitro by incubation of fresh tissue fragments with 5-bromo
2-deoxyuridine (BrdU), a thymidine analogue. Nuclei of cells in
active DNA synthesis were stained by an anti-BrdU monoclonal
antibody (Mab).
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A new procedure is described to generate
single-stranded DNA by exonuclease III (Exo III) digestion for
bromodeoxyuridine (BrdU) immunocytochemistry on tissue sections.
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Immunocytochemical detection of S-phase cells
in normal and neoplastic cervical epithelium by anti-BrdU
monoclonal antibody.
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Immunohistochemistry (IHC)
DNAse I pre-treatment markedly enhances
detection of nuclear cyclin-dependent kinase inhibitor p57Kip2
and BrdU double immunostaining in embryonic rat brain.
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BrdU immunohistochemistry for studying adult
neurogenesis: Paradigms, pitfalls, limitations, and validation.
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Evaluation of cell proliferation in rat
tissues with BrdU, PCNA, Ki-67(MIB-5) immunohistochemistry and
in situ hybridization for histone mRNA.
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Fixation in methacarn gave the highest
labeling index (16.4%), which was comparable to that observed in
unfixed frozen sections (17.5%). Formalin fixation alone
dramatically suppressed the labeling index (0.3%), which was
only partially recovered using various antigen retrieval
techniques (2.1-8.1%). Methacarn fixation is recommended for
prospective studies in which BrdU detection is planned because
of the quantitative recovery of epitope and the simplicity of
the approach.
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bromodeoxyuridine (BrdU) immunohistochemistry
in undecalcified adult rat tibiae to study cell kinetics in
various bone compartments: primary and secondary spongiosae,
periosteum, and bone marrow.
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Image analysis of bromodeoxyuridine (BrdU)
staining for measurement of S-phase in rat and mouse liver.
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volume and DNA synthesis of the
neuroepithelium in induced myeloschisis in Long-Evans rats as
shown by hematoxylin-eosin and BrdU/antiBrdU immunohistochemical
staining patterns were examined at different stages of embryonal
development.
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possibility of detecting in situ
proliferating myosatellite cells during postlarval muscle growth
in the carp (Cyprinus carpio, L.) by means of BrdU and PCNA (Cyclin)
immunohistochemistry has been evaluated on paraffin embedded
sections.
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Colonic crypt cell proliferation was measured
using a monoclonal antibody to bromodeoxyuridine by an
immunohistochemical technique.
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Immunohistochemical observations of dividing
cells in olfactory epithelium using anti-BrdU antibody.
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